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Scorpions
Scorpiones

Scorpions are eight legged venomous invertebrates belonging to the class Arachnida, and the order Scorpiones. They are related to spiders, mites, ticks, and harvestmen as well as other members of the Arachnida class. They possess an extended body and a segmented, erectile tail ending with the telson (the sting). There are roughly 1,300 species of scorpions worldwide.

Range & Habitat

Though most prolific and diverse in warm habitat, scorpions have adapted to a wide range of environments, including plains and savannahs, deciduous forests, mountainous pine forests, rain forests and caves. Scorpions have been found at elevations of over 12,000 feet in the Andes Mountains in South America and in the Himalayas of Asia, as well as the Alps. In snowy areas, they hibernate during the cold months of the year. In drought areas they may aestivate (pass the summer in a dormant or torpid state).

About 90 species are found in the U.S. All but four of these naturally occur west of the Mississippi River. Scorpions are most common in southern Arizona and in parts of Texas and central Oklahoma.

Description

The scorpion's body has two parts, a cephalothorax which contains the prosoma, or head; and the abdomen. The cephalothorax is covered by the carapace, a hard bony or chitinous outer covering. The carapace usually suports a pair of median eyes at the top center. Two to five pairs of lateral eyes are found at the front corners of the carapace, though a few cave and montane forest litter-dwelling scorpions are eyeless. Chelicerae, the scorpion's mouthparts, and a pair of pedipalps, or claws used for prey capture and mating complete the head anatomy. The pedipalps are covered with trichobothria, sensory setae, that sense air-borne vibrations.

The abdomen is made up of the mesosoma, the main body, and the metasoma, the tail. The mesosoma, protected by bony armour, contains the lungs, digestive organs and sexual organs, as well as bearing 4 pairs of walking legs and the pectines. The tips of the legs have small organs that detect vibrations in the ground. The pectines are feathery sensory organs which hang beneath the abdomen and trail on the ground. They are coated by chemosensors that provide detection of minute chemical signals that are thought to alert the scorpions to the approach of prey and also to be of use in mating behavior. The respiratory structure, known as "book lungs," are spiracles that open into the scorpion's body. The surfaces of the legs, pedipalps, and body are also covered with thicker hairs that are sensitive to direct touch.

The metasoma curves up and ends in the telson, which bears the bulbous vesicle containing the venom glands and a sharp, curved aculeus which delivers the venom.

Most species of scorpions reach adulthood at a length of between 2 and 3 inches.The longest scorpion in the world is probably the African Scorpion (Hadogenes troglodytes) which grows to over 8 inches in length. In the U.S., members of the genus Hadrurus (giant desert hairy scorpions) are probably the largest, growing to a length of about 5 inches.

Behavior

Scorpions are nocturnal. They often ambush their prey, lying in wait as they sense its approach. They consume all types of insects, spiders, centipedes, and other scorpions. Larger scorpions may feed on vertebrates, such as smaller lizards, snakes, and mice if they are able to subdue them. They capture their prey with their pedipalps, paralyzing them with their venom as well if necessary. The immobilized prey is then subjected to an acid spray that dissolves the tissues, allowing the scorpion to suck up the remains.

As well as being predators, scorpions are also prey. Many types of creatures, such as centipedes, tarantulas, insectivorous lizards, birds (especially owls), and mammals, including shrews, grasshopper mice, and bats hunt scorpions for food.

Life Cycle

The pedipalps are used in scorpion courtship behavior. The male performs a kind of dance with the female, grabbing her pedipalps with his own and dragging her across the ground until he locates a preferred place to deposit his spermatophore, which is then drawn up into the female's genital pore, near the front on the underside of her abdomen. Some species' courtships include a sexual sting of the female by the male.

Scorpion gestation periods vary from several months to a year and a half, depending on the species. Each brood will consist of about 24-35 young. They are viviparious - the young develop as embryos in the female's ovariuterus. The young scorpions are born two at at time, climbing onto their mother's back to be carried there until their first molt in about two weeks, when they will be large and strong enough to take care of themselves.

Scorpions do not metamorphasize as they grow, changing only in size and sometimes to a deeper color with each molt. Typically five or six molts over two to six years are required for the scorpion to reach maturity. The molting is accomplished by a split in the outer covering through which the scorpion must crawl in order to grow.

Scorpion lifespans range from three to five years, though some species are thought to live 10-15 years. Some kinds of scorpions show more sophisticated social behaviors, like colonial burrowing, and living in familial groups that may share burrows and food.

Scorpion Venom

Scorpion venom is used to subdue prey and to defend against threats, as well as in the mating process. The composition and action of the venom varies from species to species. The venoms are mixtures of salts, small molecules, peptides, and proteins. The peptides are specialized; some act against invertebrates and some against vertebrates, and some target both. This complex formula results in a neurotoxin which depolarizes the nervous system of the victim.

Scorpions are apparently able to regulate the delivery of the venom in scale to the size of their target. Some scorpions are known to produce a transparent prevenom in addition to the more potent opaque venom which is loaded with additional toxin. The use of the prevenom occurs at the initiation of the threat or opportunity. If the action persists, the opaque venom is released. These abilities enable the scorpion to conserve the venom for use when it is needed most, for larger predators or prey.

Medically Significant Scorpion Species

Most of the more venomous scorpions have lighter, more delicate pedipalps and larger, stronger tails. The Buthidae family contains most of the scorpions dangerous to man. They can generally be distinguished by the triangular sternal plate on their ventral side. Other species' sternal plates are more square or pentagonal.

Only one species of scorpion in North America and about 20 others worldwide have venom potent enough to be dangerous to human beings. The North American species, Centruroides exilicauda (formerly called C. sculpturatus), is found over much of Arizona and Mexico. It is also known as the Arizona Bark Scorpion. A small population occurs in extreme southeastern California, and a few records exist for southern Utah and small parts of Texas, New Mexico and Nevada. The venom of this scorpion can cause severe pain and swelling at the site of the sting, numbness, frothing at the mouth, respiratory difficulties, muscle twitching, and convulsions. The sting is more dangerous to infants, small children and the elderly. Death is rare, especially in more recent times. An antivenom was created by Arizona State University but is no longer being produced, and is not FDA approved. The FDA has recently given approval for clinical trials to evaluate a Mexican antivenom for use in the United States.

Medically significant species of scorpion occur worldwide:

  • In the Mediterranean and North Africa - Buthus, Leiurus, Androdoctonus and Leiurus
  • In Western and Southern Africa - Parabuthus
  • Across Southern Africa to Southeast Asia - Buthotus (also known as Hottentotta)
  • In Asia - Mesobuthus and the Buthotus (also known as Hottentotta)
  • South America - Tityus

    Ancestry

    Scorpions have been found in many fossil records, including coal deposits from the Carboniferous Period and in marine Silurian deposits. They are thought to have existed in some form since about 425–450 million years ago. They are believed to have an oceanic origin, with gills and a claw like appendage that enabled them to hold onto rocky shores or seaweed.


    
    
    


Freakshow, (Bob)
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